Research Index

ARA-290 Research Index

PSI surfaces 50 peer-reviewed studies on ARA-290, prioritized by evidence strength: human clinical research first, then animal studies. The full historical archive is linked below on PubMed.

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50 studies

Microenvironment-responsive injectable dynamic hydrogel for sequential antioxidant and tissue regeneration therapy of radiation-induced skin injury.

Yu X, Yang T, Bei Z et al.

Bioact Mater·2026·Animal·PubMed →

The Laccase-like Property of GHK-Cu and Its Applications in Colorimetric Sensing of Phenolic Compounds.

Chen JS, Zhu H, Chai TQ et al.

Biosensors (Basel)·2026·Animal·PubMed →

Glycyl-L-histidyl-L-lysine-Cu2(+) (GHK-Cu) Attenuates CuSO(4) or LPS induced-inflammation in Zebrafish larvae model.

Hu J, Zhang C, Wang F

Eur J Pharmacol·2026·Animal·PubMed →

Carbonless amino acids and a carbonless GHK peptide.

Skurski P, Anusiewicz I

Phys Chem Chem Phys·2026·Animal·PubMed →

Golgi-targeted copper delivery strategy via enhancing copper-dependent proteins' activity for fascia regeneration.

Wang R, Xu Y, Saiding Q et al.

J Control Release·2026·Animal·Tissue Remodeling·PubMed →

An injectable hydroxyapatite microsphere filler loaded with GHK-Cu tripeptide for anti-Inflammatory and antioxidant.

Hu D, Zhang X, Gong S et al.

Colloids Surf B Biointerfaces·2025·Animal·Anti-Inflammatory·PubMed →

Exploring the beneficial effects of GHK-Cu on an experimental model of colitis and the underlying mechanisms.

Mao S, Huang J, Li J et al.

Front Pharmacol·2025·Animal·PubMed →

Copper Complexes with New Glycyl-l-histidyl-l-lysine-Hyaluronan Conjugates Show Antioxidant Properties and Osteogenic and Angiogenic Synergistic Effects.

Greco V, Lanza V, Tomasello B et al.

Bioconjug Chem·2025·Animal·Tissue Remodeling·PubMed →

Self-Assembled Peptide-Gold Nanoparticle 1D Nanohybrids Functionalized with GHK Tripeptide for Enhanced Wound-Healing and Photothermal Therapy.

Jeon N, Kim L, Choi SG et al.

ACS Appl Mater Interfaces·2025·Animal·Wound Healing·PubMed →

Food-Derived Tripeptide-Copper Self-Healing Hydrogel for Infected Wound Healing.

Chen H, Yang P, Xue P et al.

Biomater Res·2025·Animal·PubMed →

Palmitoyl copper peptide and acetyl tyrosine complex enhances melanin production in both A375 and B16 cell lines.

Hong M, Gui Y, Xu J et al.

Biochem Biophys Res Commun·2025·Animal·PubMed →

Novel Applications of CE-ICP-MS/MS: Monitoring of Antiaging GHK-Cu Cosmetic Component Encapsulation in Liposomes.

Zajda J, Wadych E, Ogórek K et al.

Electrophoresis·2024·Animal·Skin & Cosmetic·PubMed →

The glycyl-l-histidyl-l-lysine-Cu(2+) tripeptide complex attenuates lung inflammation and fibrosis in silicosis by targeting peroxiredoxin 6.

Bian Y, Deng M, Liu J et al.

Redox Biol·2024·Animal·PubMed →

Glycyl-l-histidyl-l-lysine prevents copper- and zinc-induced protein aggregation and central nervous system cell death in vitro.

Min JH, Sarlus H, Harris RA

Metallomics·2024·Animal·Neuroprotection·PubMed →

Rigid-flexible nanocarriers loaded with active peptides for antioxidant and anti-inflammatory applications in skin.

Wang Y, Lin J, Yu Z et al.

Colloids Surf B Biointerfaces·2024·Animal·Wound Healing, Anti-Inflammatory·PubMed →

Effects of Gly-His-Lys-D-Ala Peptide on Skin Wound Regeneration Processes.

Rakhmetova KK, Mishina ES, Bobyntsev II et al.

Bull Exp Biol Med·2024·Animal·PubMed →

Behavioral and neuropathological features of Alzheimer's disease are attenuated in 5xFAD mice treated with intranasal GHK peptide.

Tucker M, Liao GY, Keely A et al.

Aging Pathobiol Ther·2024·Animal·PubMed →

Thermodynamically stable ionic liquid microemulsions pioneer pathways for topical delivery and peptide application.

Liu T, Liu Y, Zhao X et al.

Bioact Mater·2024·Animal·PubMed →

Behavioral and neuropathological features of Alzheimer's disease are attenuated in 5xFAD mice treated with intranasal GHK peptide.

Tucker M, Liao GY, Park JY et al.

bioRxiv·2023·Animal·PubMed →

Intranasal GHK peptide enhances resilience to cognitive decline in aging mice.

Tucker M, Keely A, Park JY et al.

bioRxiv·2023·Animal·PubMed →

Liposomes as Carriers of GHK-Cu Tripeptide for Cosmetic Application.

Dymek M, Olechowska K, Hąc-Wydro K et al.

Pharmaceutics·2023·Animal·PubMed →

Phenothiazine-Based Cu(II)-Selective Fluorescent Sensor: GHK-Cu Sensing Applications.

Sahu R, Yadav S, Gunturu KC et al.

J Org Chem·2023·Animal·PubMed →

Ultrasensitive and Label-Free Detection of Copper Ions by GHK-Modified Asymmetric Nanochannels.

An P, Zhang Z, Yang J et al.

Anal Chem·2023·Animal·PubMed →

Multi working mode SPR chip laboratory for high refractive index detection.

Ren Z, Liu C, Wei Y et al.

Opt Express·2023·Animal·PubMed →

Relief of ovalbumin-induced airway remodeling by the glycyl-l-histidyl-l-lysine-Cu(2+) tripeptide complex via activation of SIRT1 in airway epithelial cells.

Zhang Q, Liu J, Deng MM et al.

Biomed Pharmacother·2023·Animal·PubMed →

Improved laccase production by Trametes versicolor using Copper-Glycyl-L-Histidyl-L-Lysine as a novel and high-efficient inducer.

Wang F, Yu X, Yu Z et al.

Front Bioeng Biotechnol·2023·Animal·PubMed →

Differential Effects of Histidine and Histidinamide versus Cysteine and Cysteinamide on Copper Ion-Induced Oxidative Stress and Cytotoxicity in HaCaT Keratinocytes.

Ha JW, Choi JY, Boo YC

Antioxidants (Basel)·2023·Animal·PubMed →

Synergy of GHK-Cu and hyaluronic acid on collagen IV upregulation via fibroblast and ex-vivo skin tests.

Jiang F, Wu Y, Liu Z et al.

J Cosmet Dermatol·2023·Animal·Tissue Remodeling·PubMed →

Glycyl-l-histidyl-l-lysine-Cu(2+) rescues cigarette smoking-induced skeletal muscle dysfunction via a sirtuin 1-dependent pathway.

Deng M, Zhang Q, Yan L et al.

J Cachexia Sarcopenia Muscle·2023·Animal·Muscle Recovery·PubMed →

Glycyl-L-histidyl-L-lysine-Cu(2+) attenuates cigarette smoke-induced pulmonary emphysema and inflammation by reducing oxidative stress pathway.

Zhang Q, Yan L, Lu J et al.

Front Mol Biosci·2022·Animal·PubMed →

Biomimetic Hydrogel Scaffolds with Copper Peptide-Functionalized RADA16 Nanofiber Improve Wound Healing in Diabetes.

Yang X, Zhang Y, Huang C et al.

Macromol Biosci·2022·Animal·Wound Healing·PubMed →

Polyaspartic acid, 2-acrylamido-2-Methyl propane sulfonic acid and sodium alginate based biocompatible stimuli responsive polymer gel for controlled release of GHK-Cu peptide for wound healing.

Sharma S, Anwar MF, Dinda AK et al.

J Biomater Appl·2022·Animal·Wound Healing·PubMed →

Efficient Ternary Polymer Solar Cells with Tunable Crystallinity and Phase Separation of Active Layers via Incorporating GHK-Cu.

Huang J, Yu H

ACS Appl Mater Interfaces·2021·Animal·PubMed →

Protective and Anti-Aging Effects of 5 Cosmeceutical Peptide Mixtures on Hydrogen Peroxide-Induced Premature Senescence in Human Skin Fibroblasts.

Wu Y, Cao K, Zhang W et al.

Skin Pharmacol Physiol·2021·Animal·Tissue Remodeling, Skin & Cosmetic·PubMed →

Enhanced angiogenic effects of RGD, GHK peptides and copper (II) compositions in synthetic cryogel ECM model.

Zoughaib M, Luong D, Garifullin R et al.

Mater Sci Eng C Mater Biol Appl·2021·Animal·Wound Healing·PubMed →

Coassemble dopamine and GHK tripeptide into fluorescent nanoparticles for pH sensing.

Zheng F, Guo J, Khan AJ et al.

Luminescence·2021·Animal·Neuroprotection·PubMed →

Combination Tripeptide/Hexapeptide Serum with 1540 nm Nonablative Fractional Laser for the Treatment of Striae Distensae: A Pilot Study.

McGuinn KP, Ross NA, Wang JV et al.

Skinmed·2020·Human·PubMed →

Hydrogels based on low-methoxyl amidated citrus pectin and flaxseed gum formulated with tripeptide glycyl-l-histidyl-l-lysine improve the healing of experimental cutting wounds in rats.

Synytsya A, Poučková P, Zadinová M et al.

Int J Biol Macromol·2020·Animal·Wound Healing·PubMed →

Ternary Cu(II) Complex with GHK Peptide and Cis-Urocanic Acid as a Potential Physiologically Functional Copper Chelate.

Bossak-Ahmad K, Wiśniewska MD, Bal W et al.

Int J Mol Sci·2020·Animal·PubMed →

Expression and Purification of Recombinant GHK Tripeptides Are Able to Protect against Acute Cardiotoxicity from Exposure to Waterborne-Copper in Zebrafish.

Hsiao CD, Wu HH, Malhotra N et al.

Biomolecules·2020·Animal·PubMed →

Theoretical study of copper binding to GHK peptide.

Alshammari N, Platts JA

Comput Biol Chem·2020·Animal·PubMed →

Enhanced biological properties of collagen/chitosan-coated poly(ε-caprolactone) scaffold by surface modification with GHK-Cu peptide and 58S bioglass.

Molavi AM, Sadeghi-Avalshahr A, Nokhasteh S et al.

Prog Biomater·2020·Animal·PubMed →

The potential of GHK as an anti-aging peptide.

Dou Y, Lee A, Zhu L et al.

Aging Pathobiol Ther·2020·Animal·PubMed →

Protective effects of GHK-Cu in bleomycin-induced pulmonary fibrosis via anti-oxidative stress and anti-inflammation pathways.

Ma WH, Li M, Ma HF et al.

Life Sci·2020·Animal·Anti-Inflammatory, Tissue Remodeling·PubMed →

Effect of oligoarginine conjugation on the antiwrinkle activity and transdermal delivery of GHK peptide.

Hur GH, Han SC, Ryu AR et al.

J Pept Sci·2020·Animal·Tissue Remodeling, Skin & Cosmetic·PubMed →

In Vitro and in Vivo Studies of pH-Sensitive GHK-Cu-Incorporated Polyaspartic and Polyacrylic Acid Superabsorbent Polymer.

Sharma S, Anwar MF, Dinda A et al.

ACS Omega·2019·Animal·PubMed →

Electrophoretic deposition of GHK-Cu loaded MSN-chitosan coatings with pH-responsive release of copper and its bioactivity.

Ning C, Jiajia J, Meng L et al.

Mater Sci Eng C Mater Biol Appl·2019·Animal·Antimicrobial·PubMed →

Evaluation of the effects of topical tripeptide-copper complex and zinc oxide on open-wound healing in rabbits.

Cangul IT, Gul NY, Topal A et al.

Vet Dermatol·2006·Human·Wound Healing·PubMed →

Effects of topical copper tripeptide complex on CO2 laser-resurfaced skin.

Miller TR, Wagner JD, Baack BR et al.

Arch Facial Plast Surg·2006·Human·Wound Healing·PubMed →

A prospective randomized evaluator-blinded trial of two potential wound healing agents for the treatment of venous stasis ulcers.

Bishop JB, Phillips LG, Mustoe TA et al.

J Vasc Surg·1992·Human·Wound Healing·PubMed →